On the other 3 DSS LITE boxes are they demonstrating the same problems?

Was all the USB disks formatted as FAT or FAT32?

We have the S5000PAL motherboard (Intel SRS2500) with 1x Xeon 3.73GHz, Bios Version s5000.86b.10.00.0084 and BIOS Build Date 10/17/2007 w/ 2GB Memory FB ECC but tested with iSCSI only not with SMB. We could try but don’t think it is a OS issue for now.

Any chance you can check the logs from the GUI from Status > Hardware > Logs then click on the "Download" button (try with Firefox as IE will save this as a PHP file).

Then open and search for the log file of dmesg.log and post them. I want to see if there could be some hardware issues in the background.